Poster Session Abstracts

Effects of passive dendritic properties on the dynamics of an oscillating neuron

Michael Schwemmer, Timothy Lewis

Dendritic properties can have substantial effects on the dynamics of single neurons and the activity in neuronal networks. For instance, the architecture of the dendritic tree can alter the firing pattern of a neuronal oscillator, and dendritic filtering can change the phase-locking behavior in networks of neuronal oscillators. From a more basic standpoint, dendritic properties can affect the firing rates of neuronal oscillators in response to constant input. Even when considering passive dendrites, these effects are not as easy to understand as it might seem. That is, the dendritic "load" can have surprising effects on a neuron.s firing frequency. Kepler et al. 1990 found that coupling a somatic oscillator to a passive neuronal compartment can sometimes increase or decrease the firing frequency. However, their explanation was specific to a two variable piecewise linear relaxation oscillator. Thus, the general mechanisms by which dendritic properties affect frequency remain to be clarified.
To probe this issue further, we model a neuron as a thin passive dendritic cable attached to an isopotential somatic oscillator, i.e. a "ball and stick" model. We then use the theory of weakly perturbed oscillators to derive an equation for the change in frequency of the oscillator due to the perturbation resulting from the presence of the dendrite. Intuitively, if the reversal potential of the dendrite is lower (higher) than the average voltage of the somatic oscillations, then the firing frequency of the somatic oscillator will decrease (increase) as the radius of the dendrite increases. We show that this is indeed the case when the phase response curve of the somatic oscillator has a positive average value. However, when the average value of the phase response curve is negative, our equation shows that the intuitive prediction described above would be reversed: the firing frequency of the oscillator increases as the radius of the dendrite increases (at least initially). Curiously, for values of the dendritic reversal potential close to the average voltage of the somatic oscillations, there is a non-monotonic dependence of the firing frequency on the dendritic radius. We confirm these results using direct numerical simulations.
Our results show that the addition of a passive dendrite to a neuronal oscillator can sometimes have counter-intuitive effects on firing frequency. Furthermore, we show that these results can be understood in terms of the somatic oscillator.s phase response curve.

Modeling the Hydraulics of Root Growth in Three-Dimensions with Internal Water Sources

Brandy Wiegers, Angela Y. Cheer, Wendy K. Silk

Primary growth is characterized by cell expansion facilitated by water uptake generating hydrostatic (turgor) pressure to inflate the cell, stretching the rigid cell walls. The multiple source theory of root growth is based on the proposal that phloem and protophloem transport water into the growth zone from the mature tissue higher in the root. Here protophloem water sources are used as boundary conditions in a generalized three-dimensional model of growth sustaining water potentials in primary roots. The model predicts small radial gradients in water potential, with a significant longitudinal gradient. The results improve the agreement of theory with empirical studies for water potential in the primary growth zone of roots of Zea mays. A sensitivity analysis quantifies the functional importance of small root diameter and apical phloem differentiation in permitting growth under water stress. This presentation will complete the 4 year presentation series that I've presented for the RMA and will provide the conclusion of my thesis work.

Proteomic Aanalysis by 2-D PAGE for Response to Low Dose Ionizing Radiation and Arsenic Exposure

Dan Li, Susanne R. Berglund, Alison Santana, Zelanna Goldberg

In the field of cancer research, proteomic techniques have been applied to identify and characterize the protein signatures of human cancer cells and tissues for patient diagnosis, toxicity monitoring, and therapeutic intervention. Two-dimensional gel electrophoresis (2DGE) is a principle technique in proteomics. The objective of differential analysis of protein expression data generated by 2DGE is to identify the protein expression that is changed between biological samples taken under differing conditions. A case study on the effect to human cells of combined exposures of low dose arsenic and low dose ionizing radiation is used to present our statistical differential analysis, which includes data preprocessing, analysis of variance, adjustment for multiple comparisons and chi-square test. Several statistical methods were compared. ANOVA using empirical Bayesian estimate of protein-specific variance combined with pFDR adjustment gave the highest yield of differential protein expression while minimizing the false discovery rate.

Circle Packings and Penrose Tilings

Matthew Stamps

A circle packing is a configuration of circles with prescribed tangencies corresponding to a given triangulation. In fact, given a triangulation, there is a well-established algorithm for creating its associated circle packing. In this project, we will describe circle packings defined by triangulations that arise from Penrose tilings. Penrose tilings are interesting because, though lacking translational symmetry, they are highly ordered through a process known as inflation. This presentation will describe a gluing process which explains how the geometry of Penrose tilings is reflected in the corresponding circle packings at high levels of inflation.

Distribution Modeling of Anas platyrhynchos Under Ideal Free and Variable Conditions

Matthew Reed, Dai-Ying Wu, Andy Huang

Population dynamics can drastically affect an individual.s decision to forage for resources within a given region or relocate to an area with higher rate of net energy gain, resulting in higher fitness. Ecologists have analyzed foraging behavior using various mathematical models and computer simulations. Organisms can distribute randomly or make a conscious choice between two locations by weighing costs and benefits, with the latter potentially resulting in an ideal free distribution (IFD). The IFD model predicts that, at equilibrium, organisms will optimally distribute themselves in proportion to the net amount of resources available between a number of locations. We used a genetic algorithm (GA) simulation to predict distributions of organisms in two scenarios: one consisting of a constant return rate and the other with a variable return rate. The GA served to predict an optimal foraging strategy regarding distribution between two patches with differing rates of energy return over many generations. Experimentally, we tested our predictions against populations of mallard ducks (Anas platyrhynchos). We found that fewer ducks stayed in the high return rate patch relative to the prediction made by our GA. We hypothesize that ducks in a wild population, unlike those living within a park, as was used for our study, would exhibit distributions more consistent with our predictions.
Co-authors (in alphabetical order): Tania Gonzalez, Andy Huang, Mary Jacklin, Michelle Jensen, Christopher Mosser, Matthew Reed, Tushar Rawat, Daniel Suderow, Dai-Ying Wu
Faculty Advisors: Carole Hom Richard McElreath Andy Sih

Mathematical Modeling of Isoflurane Action on Lamprey Spinal Neurons

Tamara J Schlichter, Anne C Smith, Steven L Jinks, Timothy J Lewis

Anesthetics are believed to modulate activity of a variety of voltage-gated and ligand-gated ion channels. However, the exact biophysical mechanisms underlying anesthesia remain unclear. Recent experimental evidence suggests that the volatile anesthetic isoflurane targets the TREK and TASK two-pore potassium conductances and the persistent sodium conductance (Jinks et al. unpublished results). Furthermore, recent data indicate that volatile anesthetics produce immobility, a fundamental element of anesthesia, predominantly through direct action at the level of the spinal cord.
We use mathematical modeling and experiments on the lamprey spinal cord to study the effects of isoflurane on neuronal activity. Our experimental results on the disinhibited spinal cord preparation suggest that the excitatory interneurons of the CPG are the main target of isoflurane. As the anesthetic concentration increases, ventral root activity (assumed to be representative of the activity of spinal cord excitatory interneurons) transitions from bursting to silent. However, in some animals the activity transitions directly from bursting to silent where in others it transitions from bursting to a brief period of tonic firing before falling silent.
We incorporate the anesthetic-sensitive TREK, TASK and persistent sodium conductances into a preexisting detailed biophysical model of the lamprey excitatory interneuron, and a canonical bursting model. We then perform a thorough bifurcation analysis on these models. Our results suggest that the anesthetic effects of TREK, TASK and persistent sodium currents alone are sufficient to account for the transition from bursting to silent, but are not sufficient to account for a robust transition from bursting to tonic to silent.

High-Resolution Radar via Compressed Sensing

Matthew Herman, Thomas Strohmer

A stylized compressed sensing radar is proposed in which the time-frequency plane is discretized into an N by N grid. Assuming the number of targets K is small (i.e., K is much less than N^2), then we can transmit a sufficiently ``incoherent'' pulse and employ the techniques of compressed sensing to reconstruct the target scene. A theoretical upper bound on the sparsity $K$ is presented. Numerical simulations verify that even better performance can be achieved in practice. This novel compressed sensing approach offers great potential for better resolution over classical radar.

The Population Dynamics of a Vernal Pool Plant: The Effect of Habitat Fragmentation and Restoration on Population Persistence and Abundance

Matthew Holden, Lauren Lui, Motoki Wu, Margot Wood, Vivian Tang, Shabnam Yekta, Pak Kwong

One of the greatest challenges in conservation biology is assessing the impacts of habitat loss and fragmentation on population viability. These challenges are especially relevant to vernal pools, fragmented seasonal wetlands, because they contain many rare and endemic species. Although, vernal pools once occupied a large portion of the California landscape, increasing agricultural and suburban development has caused habitat loss and fragmentation. However, there has been no theoretical framework to guide conservation efforts. Here, we developed a mathematical model and computer simulations to describe the population dynamics of an endangered vernal pool plant. We used analytical techniques to determine general persistence conditions for the plant population and used the computer simulations to determine the effect of the pools’ spatial configuration on the plant’s population dynamics. The simulation results suggest that pool size has a larger influence on population persistence than the distance between pools. Furthermore, when the total area of pool coverage is fixed, a landscape consisting of a few large pools increases the population’s abundance and the likelihood of persistence more effectively than one with many small pools.

Multi-index Transportation Polytopes

Edward Kim, Jesứs De Loera, Shmuel Onn, Francisco Santos

Convex polyhedra are sets of feasible solutions to linear optimization problems.  The graph of a polytope is intimately connected to Dantzig's simplex family of methods for solving linear programs.  A simplex pivot rule resolves ambiguities when an iteration of the simplex method presents multiple choices.  Bounding the diameters of the graphs of polytopes is particularly interesting since the diameter of the graph is a lower bound on the number of iterations required for the simplex method using any pivot rule.  The Hirsch Conjecture asserts that the diameter of polytope has a linear upper bound.
In this poster, we consider the graphs of multi-index transportation polytpes. Transportation polytopes are well-known objects in operations research and mathematical programming.  We discuss a computer implementation to systematically enumerate all combinatorial types of non-degenerate multi-index transportation polytopes. Our exhaustive lists helped us answer some conjectures of Yemelichev et al. (1984).  We give a quadratic upper bound for the diameter of the one-skeleton graph for triply-indexed axial transportation polytopes.

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